遗迹化石Trichophy...统湄潭组中的首次报道及意义_张小乐.pdf
《遗迹化石Trichophy...统湄潭组中的首次报道及意义_张小乐.pdf》由会员分享,可在线阅读,更多相关《遗迹化石Trichophy...统湄潭组中的首次报道及意义_张小乐.pdf(12页珍藏版)》请在咨信网上搜索。
1、 收稿日期:2022-01-05;改回日期:2022-06-22;录用日期:2022-08-26*中国科学院南京地质古生物研究所古生物学和地层学国家重点实验室(20192112)、中国地质大学(武汉)生物地质与环境地质国家重点实验室(GBL22002)、中国科学院战略性先导科技专项(XDB26000000)和国家自然科学基金(42172003)联合资助。*通讯作者:张小乐,助理研究员,e-mail: 古生物学报,61(4):603614(2022年12月)Acta Palaeontologica Sinica,61(4):603614(December 2022)研究论文 DOI:10.198
2、00/ki.aps.2022001 遗迹化石Trichophycus在贵州仁怀下中奥陶统湄潭组中的首次报道及意义*张小乐1,2*张竹桐3 黄 璞1 王 怿1 吴荣昌1 1 现代古生物学和地层学国家重点实验室,中国科学院南京地质古生物研究所,南京 210008;2 生物地质与环境地质国家重点实验室,中国地质大学,武汉 430074;3 中国科学院大学,北京 100049 提要 遗迹化石研究对揭示奥陶纪生物大辐射事件期间,底栖生物在陆棚区定居的具体过程有重要意义。Trichophycus在寒武系奥陶系有广泛报道,但是针对其时空分布、演化轨迹、造迹生物属性和生态等的系统总结和讨论较少。本文首次在贵州
3、仁怀地区下中奥陶统湄潭组内陆棚沉积中报道了遗迹属Trichophycus。该遗迹化石由宽的U型、平直或弯曲的潜穴管以前进式方式构成潜穴系统。底部和侧部以纵向排列的、平行或近似平行的抓痕为特征,发育后退式蹼状构造,不发育衬里。上述特征支持将其划归到T.venosus Miller,1879。湄潭组中Trichophycus可能为一类未知的节肢动物居群所建造的觅食/居住遗迹。地质历史中Trichophycus的造迹生物来自多个类群,是行为趋同的产物。作为Glossifungites遗迹相的常见分子,它的出现可能指示沉积物的初始硬结或存在较短的沉积间断。Trichophycus相对狭窄的沉积相分布区
4、间表明它可以作为浅海固底的指相遗迹化石。作为先锋遗迹属,Trichophycus的地史分布表明,全球各板块底栖造迹生物在陆棚区的定居是不等时的。湄潭组中Trichophycus的出现,是华南扬子区奥陶纪生物大辐射事件期间,底栖造迹生物探索陆棚区生态空间的一个缩影。关键词 潜穴 “奥陶纪生物大辐射事件”古生态 古环境意义 演化轨迹 中文引用 张小乐,张竹桐,黄璞,王怿,吴荣昌,2022.遗迹化石Trichophycus在贵州仁怀下中奥陶统湄潭组中的首次报道及意义.古生物学报,61(4):603614.DOI:10.19800/ki.aps.2022001 英文引用 Zhang Xiao-le,Z
5、hang Zhu-tong,Huang Pu,Wang Yi,Wu Rong-chang,2022.First report of Trichophycus from the LowerMiddle Ordovician Meitan Formation of Renhuai area,Guizhou and its significance.Acta Palaeonto-logica Sinica,61(4):603614.DOI:10.19800/ki.aps.2022001 First report of Trichophycus from the LowerMiddle Ordovic
6、ian Meitan Formation of Renhuai area,Guizhou and its significance ZHANG Xiao-le1,2,ZHANG Zhu-tong3,HUANG Pu1,WANG Yi1,WU Rong-chang1 1 State Key Laboratory of Palaeobiology and Stratigraphy,Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences,Nanjing 210008,China;2 State Key La
7、boratory of Biogeology and Environmental Geology,China University of Geosciences,Wuhan 430074,China;3 University of Chinese Academy of Sciences,Beijing 100049,China 604 古 生 物 学 报 61 卷 Abstract Ichnofossils provide invaluable information in understanding the process of colonization of infauna in the
8、shallow-marine shelf settings during the Great Ordovician Biodiversification Event.The ichnofossil Trichophycus is widely reported from the Cambrian-Ordovician strata.However,only a few studies have been conducted to investigate its temporal and spatial occurrences,evolutionary paths,trace maker ide
9、nty as well as its ecological features.Here,we report Trichophycus from the Lower-Middle Ordovician Meitan Formation of Renhuai area,Guizhou Province for the first time.These fossils consist of relatively wide,straight to curved,U-shaped burrow segments that are arranged in a protrusive mode.The low
10、er and lateral surfaces of the burrows are characterized by sets of longitudinally arranged,parallel to sub-parallel striae.The burrow lacks lining and the infill shows retrusive spreite structures.These diagnostic characteristics allow an assignation to T.venosus Miller,1879,which is interpreted as
11、 a dwelling/feeding structure constructed by a colony of unknown arthropods,while the trace maker of reported Trichophycus elsewhere may come from various groups,representing behavioral convergence.Trichophycus venosus is a member of the Glossifungites ichnofacies,possibly indicating an initial indu
12、ration of the sea floor or a short-lived sedimentary hiatus.The ichnofossil Trichophycus may serve as a reliable indicator of firmground in shallow-marine settings.The geological distribution of Trichophycus,a pineer ichnogenus,suggests a diachronic infaunalization in shelf settings between differen
13、t blocks.Occurrence of Trichophycus in the Meitan Formation represents colonization of the shelf substrate during the Great Ordovician Biodiversification Event on the Yangtze platform,South China.Key words burrow,the Great Ordovician Biodiversification Event,paleoecology,paleoenvironmental significa
14、nce,evolutionary trail SYSTEMATIC ICHNOLOGY Ichnogenus Trichophycus Miller and Dyer,1878 Diagnosis Burrow system wide,U-or L-shaped,with hair-like scratches on the bottom and side surfaces of the tunnel;lining not developed or very thin;litholo-gy of the infill similar to that of the overlying sands
15、tone;spreite structures well-developed.Remarks The presence of scratches is highly affected by the preservation states and characteristic of the substrate.The scratches may be inconspicuous in certain circumstances.In this situation,distinguishing Trichophycus from Teichichnus Seilacher,1955,another
16、 ichnogenus whose burrows have well-developed spreite structures,could be quite tricky(Frey and Chowns,1972;Bergstrm,1976).According to Jensen(1997),Trichophycus is characterized by the common presence of vertical branches and closely arranged spreite struc-tures,while branching is seldomly observed
17、 in Teichichnus,and its spreite structures are arranged in a loose manner(Osgood,1970;Frey and Chowns,1972;Seilacher,2007;Bayet-Goll et al.,2020).The ichnoge-nus Glyphichnus Bromley and Goldring,1992 is also characterized by a U-shaped form and distinctive scratches;nevertheless,the absence of branc
18、hing and spreite structures readily differentiates Glyphichnus from Trichophycus.Trichophycus venosus Miller,1879(Figs.2-C,2-E,3-A3-G)Diagnosis Burrow system composed of wide,U-or L-shaped,straight or curved tunnels;bottom surface of the tunnel characterized by longitudinally oriented,parallel/subpa
19、rallel scratches;side surfaces of the tun-nels sometimes decorated with very fine,transversely oriented scratches;spreite structures gutter-shaped;lin-ing not developed;infill composed of sand(emended from Osgood,1970).Description Trichophycus venosus is preserved as full reliefs in thin-bedded muds
20、tone,or as hypichnial convex semireliefs in thin-bedded sandstone.The bur-row system of this ichnospecies is composed of wide,U-or L-shaped,straight or curved tunnels that are stacked in a protrusive mode.The observed maximum length of the burrow system is 27 cm.The bottom and side surfaces of the t
21、unnels are decorated with longitudinally oriented scratches.Radial scratches at the terminal end of the tunnel,as noted by Osgood(1970),are not observed.The cross section of the tunnel is oval.The diameter of the cross section ranges from 1.5 to 3.0 cm.The average diameter is 2.8 cm.Statistical anal
22、ysis based on thirty-nine fragments suggests that diameters of the cross sections display a normal distribution pattern.False branching resulted from crosscutting of the tunnels is common.True branching is inclined,with a maximum penetrating depth of 10 cm.No lining is observed throughout the studie
23、d materials.Infill of the burrow contacts sharply with the host rock.The litholo-gy of well-developed,gutter-like,retrusive spreite structures is similar to that of overlying sandstone.Remarks The ichnogenus Trichophycus was erected by Miller and Dyer in 1878 based on material from the Late Ordovici
24、an strata from the Cincinnati area,USA.Two ichnospecies,T.lanosus Miller and Dyer,1878 and T.venosus Miller,1879,were recog-nized from this area(Osgood,1970).Volk(1968)erected another ichnospecies,T.thuringicum based on 4期 张小乐等:奥陶贵州遗迹化石Trichophycus;Zhang et al.:First report of Trichophycus from the
25、Ordovician of Guizhou 605 material from the Ordovician strata from the Thuringian area,Germany.In addition,Geyer and Uchman(1995)established the fourth ichnospecies,T.tripleurum,based on material from the Cambrian strata of the Nambia desert.However,according to the original description and follow-u
- 配套讲稿:
如PPT文件的首页显示word图标,表示该PPT已包含配套word讲稿。双击word图标可打开word文档。
- 特殊限制:
部分文档作品中含有的国旗、国徽等图片,仅作为作品整体效果示例展示,禁止商用。设计者仅对作品中独创性部分享有著作权。
- 关 键 词:
- 遗迹 化石 Trichophy 湄潭 中的 首次 报道 意义 张小乐
1、咨信平台为文档C2C交易模式,即用户上传的文档直接被用户下载,收益归上传人(含作者)所有;本站仅是提供信息存储空间和展示预览,仅对用户上传内容的表现方式做保护处理,对上载内容不做任何修改或编辑。所展示的作品文档包括内容和图片全部来源于网络用户和作者上传投稿,我们不确定上传用户享有完全著作权,根据《信息网络传播权保护条例》,如果侵犯了您的版权、权益或隐私,请联系我们,核实后会尽快下架及时删除,并可随时和客服了解处理情况,尊重保护知识产权我们共同努力。
2、文档的总页数、文档格式和文档大小以系统显示为准(内容中显示的页数不一定正确),网站客服只以系统显示的页数、文件格式、文档大小作为仲裁依据,平台无法对文档的真实性、完整性、权威性、准确性、专业性及其观点立场做任何保证或承诺,下载前须认真查看,确认无误后再购买,务必慎重购买;若有违法违纪将进行移交司法处理,若涉侵权平台将进行基本处罚并下架。
3、本站所有内容均由用户上传,付费前请自行鉴别,如您付费,意味着您已接受本站规则且自行承担风险,本站不进行额外附加服务,虚拟产品一经售出概不退款(未进行购买下载可退充值款),文档一经付费(服务费)、不意味着购买了该文档的版权,仅供个人/单位学习、研究之用,不得用于商业用途,未经授权,严禁复制、发行、汇编、翻译或者网络传播等,侵权必究。
4、如你看到网页展示的文档有www.zixin.com.cn水印,是因预览和防盗链等技术需要对页面进行转换压缩成图而已,我们并不对上传的文档进行任何编辑或修改,文档下载后都不会有水印标识(原文档上传前个别存留的除外),下载后原文更清晰;试题试卷类文档,如果标题没有明确说明有答案则都视为没有答案,请知晓;PPT和DOC文档可被视为“模板”,允许上传人保留章节、目录结构的情况下删减部份的内容;PDF文档不管是原文档转换或图片扫描而得,本站不作要求视为允许,下载前自行私信或留言给上传者【自信****多点】。
5、本文档所展示的图片、画像、字体、音乐的版权可能需版权方额外授权,请谨慎使用;网站提供的党政主题相关内容(国旗、国徽、党徽--等)目的在于配合国家政策宣传,仅限个人学习分享使用,禁止用于任何广告和商用目的。
6、文档遇到问题,请及时私信或留言给本站上传会员【自信****多点】,需本站解决可联系【 微信客服】、【 QQ客服】,若有其他问题请点击或扫码反馈【 服务填表】;文档侵犯商业秘密、侵犯著作权、侵犯人身权等,请点击“【 版权申诉】”(推荐),意见反馈和侵权处理邮箱:1219186828@qq.com;也可以拔打客服电话:4008-655-100;投诉/维权电话:4009-655-100。